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成熟度: Lm ?  range ? - ? cm Max length : 30.0 cm SL 雄鱼/尚未辨别雌雄; (Ref. 56353)
Adults are found in the middle reaches of rivers during the summer; migrates downstream in autumn (Ref. 559). River bed consists of cobbles, boulders and gravel in riffles and sand and gravel in pools (Ref. 56353). Feed on fishes (Ref. 559). Juveniles migrate from the spawning ground in the sea to the middle reaches of rivers to grow (Ref. 56353).
Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino, 1984. The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)
Phylogenetic diversity index (参考文献 82804): PD50 = 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00589 (0.00281 - 0.01234), b=3.15 (2.97 - 3.33), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245).
营养阶层 (参考文献 69278): 4.2 ±0.73 se; based on food items.